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HORMONES, BRAIN, AND BEHAVIOR THIRD EDITION EDITORS-IN-CHIEF Donald W. Pfaff The Rockefeller University, New York, NY, USA Marian Joëls University Medical Center Utrecht, Utrecht, The Netherlands; and University Medical Center Groningen, Groningen, The Netherlands VOLUME 3 MOLECULAR AND CELLULAR MECHANISMS VOLUME EDITOR E. Ronald de Kloet Leiden University Medical Center, Leiden, The Netherlands AMSTERDAM • BOSTON • HEIDELBERG • LONDON • NEW YORK • OXFORD PARIS • SAN DIEGO • SAN FRANCISCO • SINGAPORE • SYDNEY • TOKYO Academic Press is an imprint of Elsevier HORMONES, BRAIN, AND BEHAVIOR